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In a 2011 study, Benoist and colleagues established that the procognitive core of Nle 1 -Ang IV resides in its three N-terminal residues (Nle-Tyr-Ile), and that C-terminally truncated tetrapeptides and tripeptides retained scopolamine-reversal activity in the Morris water maze [1]

The difference lies in the substance associated with the peptide as a salt or counterion